| 
 | BIO 432 
           Field 
            Botany Phil Ganter 301 Harned Hall 963-5782  | 
       
        | Rubus 
            idaeus, the Raspberry - Member 
            of the Rose Family, fruit is an aggregate of drupes, differ from blackberries 
            in that the core is left behind when picked, so the fruit is hollow. | 
    
  
 
The Plant  
  Families You Should Know
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Purpose 
  of the page:
 
  There are a few plant 
    families that have disproportionate impact on us, and you should know which 
    they are. We will discuss only a very limited number of important plant families.
Some Important Families:
Before working with this page, you should 
  work on the Plant Morphology page, as 
  many of the terms introduced there are used here. In addition, you should also 
  work on the Plant 
  Kingdom page to see the overall 
  organization of the plants before working with the family level here. 
We are not done with the diversity of flowering 
  plants yet but we will skip over orders of plants and go to families. Why? The 
  answer is simple: for consistency. Up until now, each division covered was composed 
  of plants so similar that you could easily place a plant into it's proper division. 
  Pines, cypresses and junipers are all easy to place in the Coniferophyta. Ferns 
  and mosses are easy to recognize, too. But flowering plants are so diverse that, 
  although they all have flowers and are easy to ID as belonging to the Magnioliophyta, 
  to leave it at that would be to ignore most of the wonderful diversity of plants. 
  Families of plants are often composed of plants similar enough to discuss as 
  a group and so, we will discuss some plant families. Some families. Some taxonomies 
  of the flowering plants have over 200 families and BIOL 4320 students have collected 
  from over 100 different families over the past few years! I want you to be familiar 
  with only a few. I choose them for one or more of three reasons: they include 
  a disproportionately large number of species, they are economically important, 
  or they are wonderful. For some, this last reason might be cause to include 
  all the families, so I will amend it to "disproportionately wonderful so 
  as to counterbalance the important goal of keeping the number of families discussed 
  to the bare minimum."
In order to keep to the criteria above, 
  the number of flowering plant families you will have to know is ten. These are 
  listed below along with a general description of the members of the family and 
  a brief summary of why the family deserved to make the listing.
  - Aceraceae 
    The Maple Family. Over 150 species. Maples are 
    tree, vines (rarely) or shrubs. We normally recognize maples by the shape 
    of their simple or compound, opposite, palmately veined leaves. Look at the 
    pictures of leaves at the Brandeis site at the end of this paragraph to see 
    some maple leaves. Very few trees in this area (perhaps Sycamore and Tulip 
    Poplar) have leaves similar, so they are easy to identify. A confusion for 
    beginning students it the name "Sycamore." Sycamore can refer to 
    either Platanus occidentalis, also called the Plant Tree, or to Acer 
    pseudoplatanus, often called Sycamore but more properly called Mock Sycamore. 
    Be careful. The flowers are usually not very showy but the dry fruits are 
    double samaras, two fruits joined together. They often rotate like a helicopter 
    when they fall from the tree and this "helicoptering" can carry 
    the seed from the mother tree. This not a very important group agriculturally 
    beyond maple syrup. The wood is important in furniture manufacture. The reason 
    we include it here is that it is an important member of our forest and you 
    are almost sure to include one or more maples in your collection. More on 
    maples can be found at the Wikipedia 
    site, the Digital 
    Flowers database of the U. of Illinois, and at the DELTA 
    site (see bottom of page for more on DELTA). There are some shots of maple 
    leaves at Brandeis 
    U. Biology.
- Cactaceae 
    The Cactus Family. Over 2000 species. These are stem succulents because they 
    store water in their stems. Except for a very few species in a single genus, 
    they also have no mature leaves (sometimes new growth will briefly have succulent 
    leaves attached), although some of the stems may be flattened and look a little 
    like leaves (think of the Christmas cactus sold during the holidays). The 
    nodes produce spines instead of leaves and their anatomy is unusual enough 
    that they get the name of aureoles. They are included here because they are 
    one of only two families found only in North and South America and much of 
    my research has involved things that live in cacti. Humans have used cacti 
    as food for both use and for livestock. The fruits are most often eaten but 
    the seeds can be roasted and ground into flour. Some prickly pear pads are 
    eaten after being cleaned of spines and outer cuticle (try a nopal salad next 
    time you are in a good Mexican restaurant). As livestock feed in dry areas, 
    cacti can provide both only food and water at the same time. Some cactus flesh 
    contains toxins (alkaloids and other lipids) and these have been exploited 
    as poisons to kill fish and as medicines (as emetics or laxatives). Some of 
    the toxins are hallucinogenic. There are excellent websites on cacti at Wikipedia 
    , at the DELTA 
    taxonomy site (see bottom of page for more on the DELTA project), and at Botany-Online. 
    There are pictures of cacti at desert 
    tropicals and at a page by Mark 
    Olson. There are beautiful pictures of cactus flowers at the On 
    Closer Inspection site. Take a look at these both for their beauty and 
    to see that they all have the same basic structure. Once you have really looked 
    at a cactus blossom, you can never fail to recognize them.
- Compositae 
    The Composite or Daisy Family. Over 20,000 species in over 900 genera. The 
    members of this large family are most often herbs, although there are a few 
    bushes and trees in the family (no trees or bushes are native here). The leaves 
    vary a lot and can be either simple of compound, alternate or whorled. The 
    best way to recognize a member of the Compositae is to see its flower, which 
    is always many flowers fused together into a flower head. Each flower (called 
    a floret) sits on the fused receptacle, which is often covered by bracts. 
    The flowers may be tiny but the flower head can be large (sunflowers belong 
    to this family). The flowers along the periphery of the flower head fuse their 
    five petals into a strap-like petal that sticks out from the flower head. 
    These are called rays and the florets that produce them ray florets. The more 
    central florets usually produce very short tube corollas. The Compositae is 
    included here due to the large size of the family. In addition, although we 
    eat rather a small number of species out of this large family (artichokes, 
    lettuce, endive, radicchio, Jerusalem artichoke, and sunflower seeds) we have 
    cultivated many ornamentals from the family (marigold, zinnia, chrysanthemum, 
    dahlia, and many others). Good sites are found at 
    Botany-Online, Oregon 
    State U. (called the Asteraceae here), Wildflowers 
    and Weeds (which has lots of good pictures), and Wikipedia. 
    
- Curcurbitaceae 
    The Gourd or Melon Family. Over 800 species. Most melons are annual vines 
    with palmately-veined, simple, opposite leaves with large lobes and lots of 
    hairs stiffened into prickles. The solitary flowers are large with 5 parts 
    and often separate male and female flowers are found on the same plant. The 
    fruit is a pepo or more commonly, a melon. The family is important agriculturally. 
    Squash, zucchini, cantaloupes, honeydews, cucumbers, luffas, gourds, pumpkins, 
    hubbards, butternut squash, acorn squash, calabash, and watermelons are all 
    members of the family. They often produce triterpenoid cucurbitacins, bitter-tasting 
    oils that give the fruits a bad taste. More on melons can be found at the 
    Wikipedia site, the 
    Digital 
    Flowers database of the U. of Illinois, Biology 
    Online, and at the DELTA 
    site (see bottom of page for more on DELTA). 
- Fabaceae 
    The Pea Family. Over 18,000 species. Members of this family are also called 
    legumes. In fact, the family used to be called the Leguminosae and some still 
    use the term. As an additional bit of confusion, some taxonomists have elevated 
    some of the subgroups within the family up to family status, so Mimosa trees 
    are in the Fabaceae in some classifications and and in their own family, the 
    Mimosaceae. We will use the more inclusive form. Members of the family are 
    everything from herbs to trees. The leaves are usually alternate and pinnately 
    compound (palmately compound and simple leaves are also found but not very 
    commonly). There are usually stipules at the 
    base of the petioles. The flowers are most often perfect with a single, simple 
    ovary. It is this ovary that develops into the fruit type all Fabaceae have 
    in common: the legume. This is a pea pod. Some pods do not dehisce (split 
    open) as they dry out but most do and they split along both sides of the pod. 
    The family is important in more than one way. 
      It is perhaps the second most important family agriculturally. Peas, beans 
      of all sorts (and there are many types of beans from those whose pod is 
      edible to those whose seeds are edible), tamarind, carob, lentils, licorice, 
      and peanuts. Peanuts are unusual in that the stem with the flower grows 
      downward after fertilization and the fruits develop underground (many in 
      the South call them ground nuts). Beans are important sources of proteins 
      for those who either forgo animal proteins or cannot obtain animal protein. 
      Soy beans are particularly high in protein and have been adopted world-wide 
      as animal feed. The family includes many members that nodulate (see link 
      to a pdf file if this is confusing - just read the introduction pages) and 
      so growing them can increase the level of biologically useful nitrogen in 
      the soil. This is often a nutrient in very limited supply and so soil fertility 
      usually increases if clover or alfalfa are grown on it. The clover and alfalfa 
      are then harvested as winter forage for cattle. Kudzu is a member of the 
      family brought to the US to control erosion on hillsides. It has become 
      a problem is some areas by overgrowing trees but it does have edible fruits 
      and the leaves are good animal forage. Finally, many ornamentals belong 
      to the family, from flowers to trees. More on the Fabaceae can be found 
      at the Wikipedia site, 
      the Digital 
      Flowers database of the U. of Illinois, Reed 
      College, and at the DELTA 
      site (see bottom of page for more on DELTA) uses the name Leguminosae. 
- Fagaceae 
    The Oak or Beech Family. Over 1000 species. This is a family of trees, including 
    the dominant trees in out forests, and a few shrubs. The trees are mostly 
    deciduous but some evergreens are found in the family. The leaves are simple 
    and alternate. Stipules are usually present (for some pictures of leaves from 
    the family go to this link). 
    The flowers are often less than spectacular (an exception is the European 
    chestnut). The fruits are nuts surrounded by a cupule, which develops from 
    the involucre (a set of bracts that surrounds a flower or flower head). The 
    cupule is the acorn's cup or the husk around a hickory nut. The economic importance 
    of the family comes from the nuts produced by chestnut trees and by the valuable 
    hardwood lumber cut from many of the species. We are interested in the family 
    because they are the dominant trees in local forests and you are sure to have 
    a few members of the family in your collection. The family includes the beeches, 
    oaks, and chinquapins. More on the family can be found at the Wikipedia 
    site, the Digital 
    Flowers database of the U. of Illinois, in an online article by Henrietta 
    Chambers in the journal Oregon 
    Flora Newsletter, and at the DELTA 
    site (see bottom of page for more on DELTA).
- Orchidaceae 
    The Orchid Family. Over 25,000 species (over 100,000 hybrids and cultivars 
    are known). This is the largest family of plants. Many species have flowers 
    that are capable of pollination by one or two species of insect only and it 
    is thought that the tendency to form these specific pollination mutualisms 
    have given rise to so many species of orchids. The plants are herbs or vines 
    (vines are also called lianas). Most are terrestrial 
    but many are epiphytes growing on other plants 
    (not parasites but just using the host as a way to get more sunlight). This 
    is a monocot family and the leaves tend to be strap-like with parallel veination. 
    The flowers are zygomorphous (bilaterally symmetric) and can be quite complex. 
    An easy-to-see feature is the lip or labellum, a single tounge-like or bag-like 
    structure that comes from the center of the flower. Interest in orchids stems 
    more from a sense of beauty that from hunger. Orchids do give us an important 
    spice, vanilla. Some orchids have tubers at the 
    base of the leaves and one species in Turkey is the basis for a popular kind 
    of ice cream. Orchid flowers are prized world-wide and have lead to intense 
    collecting and breeding. When humans transfer pollen rather than insects, 
    hybrids form readily and flower breeders have exploited this tendency to produce 
    more hybrids than there are species! More information on orchids can be found 
    at Botany-Online, 
    at Wikipedia, at Reed 
    College, and at the Delta database (see bottom of page for more on the 
    delta system) page on orchids.
- Poaceae 
    The Grass Family. Between 8000 and 10,000 species. 
    Grass family members are herbs, 
    except for bamboo, which is tall but still a herb. Grasses, although found 
    in all habitats from deserts to rain forests, dominate where there is some 
    rainfall, often strongly seasonal, but not enough to support forests (they 
    are so dominant we call this biome the grassland). Many grasses employ the 
    C4 
    pathway, which reduces photorespiration and water loss through transpiration 
    (to see why, follow this link). 
    This is one of a suite of adaptations to low-water stress conditions. Many 
    plants spread through underground rhizomes or above ground stolons. The stems 
    have hollow internodes (the areas between the nodes) and are often hardened 
    by the deposition of silica (SiO2, 
    glass). This anti-herbivore adaptation is also found in the leaves, which 
    can give the edge of a leaf the sharpness and strength to cut fingers, so 
    watch out. The leaves have no petioles and the leaf base attaches in a strip 
    around the node. The meristem (growing region) of the leaf is at the base 
    instead of at the tip (as it is in the vast majority of plants). This is an 
    adaptation to grazing, where the top of the leaf can be removed all at once 
    but the leaf can regrow from the base. The blade of the leaf often forms a 
    sheath that surrounds the stem before it splits and the flat, strap-like leaf 
    bends away from the stem. If you look closely at the base, there is often 
    a small extension from the base called a ligule. 
    This is a monocot family and grass leaves are classically parallel veined. 
    The flowers are small and wind-pollinated, so they tend to be inconspicuous. 
    Petals have been lost and each floret is surrounded by bracts (each of which 
    have separate names we won't use here). The fruits form on the floret spikelets, 
    which gives them the look we all know from wheat or corn ears. 
    It is almost impossible to overstate the importance 
      of the Poaceae to both our everyday lives and our history. We humans gave 
      up the hunter-gatherer lifestyle and settled into permanent communities 
      when we began to cultivate crops. Where ever civilizations grew, the staple 
      crop was almost always a grass (potatoes in the mountains of South America 
      are an exception). We sustain a population of billions mostly on grass crops 
      (or meat raised on grasses). The Poaceae contains most of our staple food 
      crops: wheat, corn & rice. In addition, beer is brewed from barley (which 
      is also eaten), and sugar can be processed from sugar cane or sorghum (also 
      called milo), although beets are also an important source of sugar. Grasses 
      are important in preserving soils with their tight network of fibrous roots 
      (see the Vetiver page 
      for a new application of this technology). Grasses are also important ornamentals 
      and might be in your collection. More on the grasses at Wikipedia, 
      a slightly diffferent version of the Wikipedia page at Biocrawler, 
      at the What-you-need-to-know-about 
      page, at Oregon 
      State U., and a page that compares monocot families with a brief description 
      of grasses and links to grass pictures at the U. 
      of Tasmania. There is a pdf that describes the grass family available 
      for downloading at the U. 
      of Texas. 
- Rosaceae 
    The Rose Family. Over 3000 species. Members of this family can be herbs, shrubs, 
    vines or, very often, trees. The leaves are usually alternately placed, can 
    be compound or simple, and usually stipules are present at the base of the 
    petiole. The flowers are 5 part or multiples of 5, solitary, with little fusion 
    of petals and lots of stamens. The fruits are most often drupes but pomes 
    are common and some achenes (the fruit is a seed 
    covered with some dried layers of tissue called the pericarp) are found in 
    the family. It is the fruits that make this family importance to us. Cherries, 
    plums, and peaches are all drupes. Sometimes drupes are aggregated into a 
    single structure as in raspberries, blackberries, black raspberries, dewberries, 
    loganberries, cloudberries, salmonberries, etc. Almonds are a nut crop from 
    a member of the family. Strawberries are rose family members with achenes 
    as fruits (they are the little black "seeds" on the outside are 
    the strawberry). What we eat as the strawberry is botanically called the accessory 
    fruit. Of course, roses are among the most popular ornamental plants in temperate 
    zones. More information on the rose family can be found at Botany-Online, 
    at Wikipedia, at 
    Reed College, and at the Delta database (see bottom of page for more on 
    the delta system) page on the Rose 
    Family.
- Solanaceae 
    The Nightshade or Potato Family. Over 2500 species. These are mostly herbs 
    (a few are vines or small trees) with simple leaves, often with hairs or prickles 
    on them, arranged alternately (may appear opposite near flowers). The flowers 
    parts are usually in fives with the petals and sepals each fused at the base. 
    The petals are often completely fused into a tube-like corolla and the ovary 
    is inferior. The fruits are either fleshy berries (tomatoes) or dry capsules 
    (peppers) that split open when totally dry. The family has a long history 
    of uses for humans, which is why it qualifies as one of the ten families. 
    The potatoes, tomatoes, eggplant, peppers, chilies, paprika, mandrake, and 
    tomatillo are all members of the family. Many family members produce alkaloids, 
    secondary chemicals found in many plant families. The alkaloids in the Solanaceae 
    often have toxic or pharmacologic effects. Jimson weed (loco weed) and deadly 
    nightshade (belladonna) can kill if too much is ingested. Even the green portion 
    of potatoes is toxic, although we usually don't eat enough to affect us. Belladonna 
    is used to dilate eyes by ophthalmologists. Tobacco is a member of the family 
    and nicotine is the alkaloid it produces. Finally, the petunia is a member 
    of the family and what would spring in Tennessee be without gardeners setting 
    out petunias? Good websites on the family are at Botanical-Online, 
    Oregon State 
    U. , Wikipedia, 
    and the site maintained by J. A. M. van 
    Balken, a man dedicated to the Solanaceae. You can see a flower with five 
    stamens and a fused corolla on the main page of his website.
There is an interesting online plant key system 
  for families based on DELTA (DEscription Language for TAxonomy) developed by 
  MJ Dallwitz of Australia's CSIRO (Commonwealth Scientific and Industrial Research 
  Organization) and colleagues. It requires that you download a small program 
  (for Windows only) which accesses an online database so that the information 
  is always up-to-date. You can get the program and instructions at the DELTA 
  homepage.
Last 
  updated July 4, 2006